John Werner, Ph.D.

John S. Werner, Ph.D.




  • Distinguished Professor


Lawrence J. Ellison Ambulatory Care Center

4860 Y St., Suite 2400
Sacramento, CA 95817
Driving Directions

Primary Phone:

Additional Phone Numbers

Physician Referrals: 800-4-UCDAVIS (800-482-3284)



Clinical Interests

Dr. Werner studies the neurophysiological computations and mechanisms that mediate human vision, particularly changes in color and spatial vision across the life span and in association with retinal and optic nerve disease.

His work has demonstrated adaptive modifications of the visual system response to changes in signals due to aging of the eye's optics and retina. His laboratory has developed state-of-the-art methods for imaging the human retina of the living eye at a cellular scale. 



Center/Program Affiliation

Eye Center


Ph.D., Psychology, Brown University, Providence RI 1979

M.A., Human Development, University of Kansas, Lawrence KS 1974

B.A., University of Kansas, Lawrence KS 1974


Instituut voor Zintuigfysiologie - TNO (Institute for Perception), Soesterberg, The Netherlands 1979

Universitat Freiburg, Abteilung fur Neurophysiologie, Freiburg, i, Br., Germany 1981

Professional Memberships

American Association for the Advancement of Science

Association for Research in Vision and Ophthalmology

International Colour Vision Society

Optical Society of America

Honors and Awards

International Colour Vision Society, Verriest Medal, 2015

Optical Society of America, Fellow�s International Lecture Award, 2012

R&D 100 Award, 2010

Lighthouse International (New York), Pisart Vision Award, 2008

Jules and Doris Stein Research to Prevent Blindness Professorship, 2000

University of Colorado, Ninetieth Annual Distinguished Research Lecture, 1998

Humboldt Research Prize for Senior Scientists in the Natural Sciences, Bonn, Germany, 1994

Select Recent Publications

Werner, J.S. & Chalupa, L.M. (Eds.) The New Visual Neurosciences. Cambridge, MA: MIT Press. 2014.

Werner JS. The Verriest Lecture: Short-wave-sensitive cone pathways across the life span. J Opt Soc Am A Opt Image Sci Vis. 2016 Mar 1;33(3):A104-22.

Schwartz DM, Fingler J, Kim DY, Zawadzki RJ, Morse LS, Park SS, Fraser SE, Werner JS. Phase-variance optical coherence tomography: a technique for noninvasive angiography. Ophthalmology. 2014 Jan;121(1):180-7.

Kim DY, Fingler J, Zawadzki RJ, Park SS, Morse LS, Schwartz DM, Fraser SE, Werner JS. Optical imaging of the chorioretinal vasculature in the living human eye. Proc Natl Acad Sci U S A. 2013 Aug 27;110(35):14354-9.

Panorgias A, Zawadzki RJ, Capps AG, Hunter AA, Morse LS, Werner JS. Multimodal assessment of microscopic morphology and retinal function in patients with geographic atrophy. Invest Ophthalmol Vis Sci. 2013 Jun 26;54(6):4372-84.

Shinomori K, Werner JS. Aging of human short-wave cone pathways. Proc Natl Acad Sci U S A. 2012 Aug 14;109(33):13422-7.